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The excitatory and inhibitory areas of the receptive field of the cortical cell balance each other, and as a result such a cell does not respond to the uniform illumination of the entire receptive field. It can only be aroused by stripes (light or dark) or contrasting with appropriately-arranged and arranged borders. 184

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ny. There are many different subgroups of such cells, and each subgroup has its own preferred orientation of the linear stimulus to which the cells respond vigorously.

 

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The question of what kind of information these cortical neurons encode remains unresolved. However, it seems clear that cells with such properties prepare the brain to perceive contrasts and to analyze forms. Some cases of contrast between adjacent areas are shown in Fig. 5: The disk looks brighter or darker, depending on which direction the lightness changes at its boundary. Those afferent ‘neurons, whose concentric receptive fields fit entirely into evenly illuminated areas of the retina, react in exactly the same way in both cases. Those cortical cells, the oriented receptive fields of which appeared on the same sites, do not respond at all. But if the oriented field falls on the dividing line, then the corresponding cortical neuron reacts as shown in Fig. 5. Thus, it can be assumed that the contrast of perceived lightness of the entire disk is due to extrapolation of the effect created by the edge neurons over its entire area. If this is so, then the creation of oppositely directed lightness drops on opposite edges of the disk would limit the visible contrast to one half of the disk.

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